Please answer a question about the following article about Aspirated consonant:

Many Indo-Aryan languages have aspirated stops. Sanskrit, Hindi, Bengali, Marathi, and Gujarati have a four-way distinction in stops: voiceless, aspirated, voiced, and breathy-voiced or voiced aspirated, such as /p pʰ b bʱ/. Punjabi has lost breathy-voiced consonants, which resulted in a tone system, and therefore has a distinction between voiceless, aspirated, and voiced: /p pʰ b/.

Which distinction has Punjabi lost?
breathy-voiced consonants