When a Teletype 33 ASR equipped with the automatic paper tape reader received a Control-S (XOFF, an abbreviation for transmit off), it caused the tape reader to stop; receiving Control-Q (XON, "transmit on") caused the tape reader to resume. This technique became adopted by several early computer operating systems as a "handshaking" signal warning a sender to stop transmission because of impending overflow; it persists to this day in many systems as a manual output control technique. On some systems Control-S retains its meaning but Control-Q is replaced by a second Control-S to resume output. The 33 ASR also could be configured to employ Control-R (DC2) and Control-T (DC4) to start and stop the tape punch; on some units equipped with this function, the corresponding control character lettering on the keycap above the letter was TAPE and TAPE respectively.

What casued the automatic paper tape reader to stop?