Sentient beings always suffer throughout saṃsāra until they free themselves from this suffering (dukkha) by attaining Nirvana. Then the absence of the first Nidāna—ignorance—leads to the absence of the others.
The absence of ignorance leads to what?
the absence of the others

The convention followed by many British publishers (including scientific publishers, like Nature, magazines, like The Economist and New Scientist, and newspapers, like The Guardian and The Times) and U.S. newspapers is to use sentence-style capitalisation in headlines, where capitalisation follows the same rules that apply for sentences. This convention is usually called sentence case. It may also be applied to publication titles, especially in bibliographic references and library catalogues. Examples of global publishers whose English-language house styles prescribe sentence-case titles and headings include the International Organization for Standardization.
What is the basis of capitalization standards for the sentence-style capitalization?
sentences

In 1898, Bell experimented with tetrahedral box kites and wings constructed of multiple compound tetrahedral kites covered in maroon silk.[N 23] The tetrahedral wings were named Cygnet I, II and III, and were flown both unmanned and manned (Cygnet I crashed during a flight carrying Selfridge) in the period from 1907–1912. Some of Bell's kites are on display at the Alexander Graham Bell National Historic Site.
Who was on Cygnet I when it crashed?
Selfridge